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Size-fractionated extract spiking

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Allelopathy (size-fractionated extract spiking)
Approach: size-fractionated culture extracts added to target species; growth or Fv/Fm response measured
Context: lab
Spatial scale: culture flask (point)
Temporal scale: hours to days
Units: cells L-1 d-1; Fv/Fm
Community captured: individual target species
Co-measurements: temperature, PAR; sometimes pH and salinity

Method Overview

Cell-free extracts or filtrates from a potentially allelopathic species are size-fractionated by molecular weight cutoff ultrafiltration membranes (e.g., < 1 kDa, 1–10 kDa, 10–100 kDa, > 100 kDa) or by reverse-phase solid-phase extraction (SPE) into polarity fractions. Each fraction is then added separately to cultures of a target species. The effect of each fraction on target species growth rate or photosynthetic efficiency (Fv/Fm) is quantified relative to solvent controls[1]. This approach identifies the molecular size range (and sometimes polarity class) of the allelopathic compound(s), aiding in their chemical characterization.

Scale of measurement

Laboratory culture; no direct in situ spatial scale. Hours to days incubation per fraction.

Data generated

Growth rate or Fv/Fm of the target species in response to each size fraction. The fraction(s) causing significant inhibition narrow down the molecular size class of the allelopathic compound.

Units & currency

Units are cells L-1 d-1 (growth) or Fv/Fm. The currency is cell abundance or photon relaxation.

Sample size

Typical samples are < 1 L in volume per fraction.

Repositories & databases

Limitations

The size fractionation assumes that the allelopathic compound is in a specific size class and that fractionation does not alter its activity (e.g., through pH changes during ultrafiltration or solid phase extraction). Compounds may interact synergistically across fractions, making single-fraction bioassays incomplete. The assumption that a specific compound or fraction exerts the allelopathic effect must be verified by chemical identification and dose-response experiments.

Example Applications & Protocols

Classic examples

  • Wu et al. (2010) Allelopathic control of cyanobacterial blooms by periphyton biofilms [1]

Recent applications

Common calculations/conversions

  • % inhibition per fraction = (Fv/Fmcontrol − Fv/Fmfraction) / Fv/Fmcontrol × 100; highest inhibition identifies the active size class.

References

  1. 1.0 1.1 Wu, J.-T., Chiang, Y.-R., Huang, W.-Y., & Jane, W.-N. (2010). Allelopathic control of cyanobacterial blooms by periphyton biofilms. Environmental Microbiology, 13(3), 604–615. https://doi.org/10.1111/j.1462-2920.2010.02363.x